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Strümpher, Werner P, Sole, Catherine L, Villet, Martin H, Scholtz, Clarke H. Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. 2014.
Strümpher, Werner P, Sole, Catherine L, Villet, Martin H, Scholtz, Clarke H. Brill; 2016. Allopatric speciation in the flightless Phoberus capensis (Coleoptera: Trogidae) group, with description of two new species.
Strümpher, Werner P, Villet, Martin H, Sole, Catherine L, Scholtz, Clarke H. Brill; 2016. Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera Scarabaeoidea)..